Abkouh ASK, Iranbaksh I, Haji Moniri M and Asri Y. 2022. New records of five Staurothele species (Verrucariaceae, Ascomycota) from Iran. – Acta Biologica Szegediensis 66: 57–61. https://doi.org/10.14232/abs.2022.1.57-61
Baradaran B, Saadatmand S, Haji Moniri M and Asri Y. 2020. New lichen records from north-east of Iran. – Iran. J. Bot. 26: 166–171. https://doi.org/10.22092/ijb.2020.351116.1293
Baradaran B, Saadatmand S, Haji Moniri M and Asri Y. 2023. New records of crustose lichen species for Iran and Asia. – Acta Biologica Szegediensis 67: 55–61. https://doi.org/10.14232/abs.2023.1.55-61
Borodin AM, Bannikov AGand SokolovV. (eds.) 1978. Red Data Book of USSR: Rare and Endangered Species of Animals and Plants. Vol. 2. – Lesnaya Promyshlennost Publishers, Moscow.
Brummitt RK and Powell CE. 1992. Authors of Plant Names. – Royal Botanic Gardens, Kew.
CABI Bioscience, CBS and Landcare Research 2009. Index Fungorum (www.indexfungorum.org) accessed Tuesday, July 8, 2025
Chakimov LA, Bronnikova VK, Matskevitch NVand Tolpyscheva TY. 1986. Geographical distribution of the rare species of lichen Aspicilia oxneriana Blum. – Bulletin of Moscow State University, Series 16, Biology 4: 8–15.
Crous PW, Gams W, Stalpers J.A, Robert V and Stegehuis G. 2004. MycoBank: an online initiative to launch mycology into the 21st century. – Studies in Mycology 50: 19–22.
Dobson FS. 2011. Lichens – An Illustrated Guide to the British and Irish Species. – Richmond Publishing Co., Richmond.
Dzhuraeva Z. 1978. Lichen flora of Central Kopet-Dag. – Ylum, Ashgabat, Turkmenistan.
Gholamhosseinian A, Sepehr A. and Emadodin I. 2020. The effects of biocrusts on soil parameters in a semi-arid pediment at north-eastern Iran. – Revista de Geomorfologie 22: 5–19. https://doi.org/10.21094/rg.2020.094
Gholamhosseinian A, Sepehr A, Sohrabi M and Emadodin I. 2021. Assessing the role of lichens in the prevention of dust emission in dryland: case study at north-eastern Iran. – Aeolian Research 50: 100697.
https://doi.org/10.1016/j.aeolia.2021.100697
Haji Moniri M. 2005. Lichen genus new to Iran. – In: Singh, J.S. et al. (eds.), Abstracts, 3rd International Conference on Plants and Environmental Pollution (ICPEP-3), Lucknow, India, 29 November – 2 December 2005, p. 35. University Press, Lucknow.
Haji Moniri M. 2009. History of lichenology in Iran, with some additional lichens from Golestan Province (N. Iran). – Iranian Journal of Botany 15: 159–163.
Haji Moniri M. 2010. A small lichen collection from Darkesh, NE Iran. – Journal of Environmental Science and Engineering 4: 36–39.
Haji Moniri M. 2014. Further investigations on
Rhizocarpon of north-eastern Iran: R. geographicum. – Journal of Mycology 2 (ID 528041): 1–5.
https://doi.org/10.1155/2014/528041
Haji Moniri M. 2018. Preliminary investigation of the ecological interaction between corticolous lichens and Juniperus excelsa (case study of the Tandoureh forests). – Journal of Plant Research 31: 54–62.
Haji Moniri M. 2025. Comprehensive field guide, identification and diversity of lichens: lichens of northeastern Iran. Islamic Azad University, Mashhad Branch, Mashhad, Iran.
Haji Moniri M and Kamyabi S. 2010. Morphological and anatomical investigation of Rhizocarpon disporum and its distribution in Razavi Khorasan Province. – Journal of Plant Science Researches 17: 36–43.
Haji Moniri M. and Kamyabi S. 2011. On Rhizocarpon viridiatrum (Wulfen) Körb. in north-eastern Iran. – Iranian Journal of Botany 17: 276–281.
Haji Moniri M,- Kukwa M. 2009. Additions to the lichen biota of Iran. – Mycotaxon 110: 155–161.
https://doi.org/10.5248/110.155
Haji Moniri M,Seaward MRD. 2005. Teloschistes, a lichen genus new to Iran. In: Singh et al. (eds), Abstracts, 3rd International Conference on Plants and Environmental Pollution (ICPEP-3), Lucknow, India, 29 November–2 December 2005, p. 144. University Press, Lucknow.
Haji Moniri M, Sipman HJM. 2009. Lichens of two nature reserves in NE Iran. – Willdenowia 39: 199–202.
https://doi.org/10.3372/wi.39.39121
Haji Moniri M,Sipman HJM. 2011. Lichens from three mountain sites in Khorasan provinces, Iran, including four species new to Iran. – Cryptogamie, Mycology 32: 145–150. https://doi.org/10.7872/crym.v32.iss2.2011.145
Haji Moniri M, Sipman HJM. 2017. New and noteworthy lichens from the semi-desert areas of North East Iran. – Iranian Journal of Botany 23: 140–144.
Haji Moniri M, Massoumi M and Fallahian F. 2005. Lichens from the Khorasan Province, Iran. – Folia Cryptogamica Estonica 41: 55–58.
Haji Moniri M, Kamyabi S and Fryday AM. 2009a. Rhizocarpon saurinum new to Asia, and other reports of Rhizocarpon species from Razavi Khorasan Province, Iran. – Mycologica Balcanica 6: 89–92.
https://doi.org/10.5281/zenodo.2549930
Haji Moniri M, Soltani A and Kamyabi S. 2009b. Some lichens from Kashmar, NE Iran. – Journal of Applied and Natural Science 1: 286–290. https://doi.org/10.31018/jans.v1i2.73
Haji Moniri M, Kamyabi S and Clayden SR. 2010. A preliminary study of Rhizocarpon macrosporum in Razavi Khorasan province (NE Iran). – Iranian Journal of Botany 16: 185–189.
Haji Moniri M, Jandaghi M and Masroornia M. 2011a. A note on lichens in the vicinity of Mashhad (Razavi Khorasan, NE Iran). – Iranian Journal of Botany 17: 133–136.
Haji Moniri M, Noferesti N, Mirasgari SJ and Tavakoli M. 2011b. Lichenized fungi of the Binaloud Mountains, NE Iran. – Mycologia Balcanica 8: 93–96. https://doi.org/10.5281/zenodo.2550428
Haji Moniri M, Sipman HJM and Schultz M. 2014. New records of lichenized and lichenicolous fungi from northeastern Iran. – Herzogia 27: 367–376. https://doi.org/10.13158/heia.27.2.2014.367
Haji Moniri M, Gromakova AB, Lőkös L and Kondratyuk SY. 2017. New members of the Megasporaceae (Pertusariales, lichen-forming Ascomycota): Megaspora iranica spec. nova and Oxneriaria gen. nova. – Acta Botanica Hungarica 59(3–4): 343–370. https://doi.org/10.1556/034.59.2017.3-4.5
Hassanzadeh Bashtian M, Sepehr A, Bahreini M and Farzam M. 2019. Microgeomorphology related soil characteristics determine the heterogeneity of biological soil crust communities. – Geografia Fisica e Dinamica Quaternaria 42: 99–112.
https://doi.org/10.4461/GFDQ.2019.42.8
Kamyabi AS, Iranbaksh A, Haji Moniri M and Asri Y. 2020. Statistical comparison of morphological and anatomical characters from Verrucaria nigrescens Pers. based on the substrate along with insects attraction in the northeast zone of Iran. – International Journal of Entomology Research 5: 115–124.
Kamyabi AS, Iranbaksh A, Haji Moniri M and Asri Y. 2023. New records of five Verrucaria species (Verrucariaceae, Ascomycota) from Iran. – Journal of Botanical Taxonomy and Geobotany 134: 191–199.
https://doi.org/10.1002/fedr.202200017
Kondratyuk SY, Jeong MH, Yu N-N, Karnefelt I, Thell A, Elix JA, Kim J, Kondratiuk AS and Hur J-S. 2013a. Four new genera of teloschistoid lichens (Teloschistaceae, Ascomycota) based on molecular phylogeny. – Acta Botanica Hungarica 55: 251–274. https://doi.org/10.1556/ABot.55.2013.3-4.8
Kondratyuk SY, Lőkös L, Zarei-Darki B, Haji Moniri M, Tchabanenko S, Galanina I, Yakovchenko L, Hooshmand F, Ezhkin AK and Hur J-S. 2013b. Five new Caloplaca species (Teloschistaceae, Ascomycota) from Asia. – Acta Botanica Hungarica 55: 41–60. https://doi.org/10.1556/ABot.55.2013.1-2.4
Kondratyuk SY, Elix JA, Kärnefelt I and Thell A. 2013c. Five new Caloplaca species (Teloschistaceae, Ascomycota) from Australia. – Folia Cryptogamica Estonica 50: 41–48. https://doi.org/10.12697/fce.2013.50.06
Kondratyuk SY, Jeong M-H, Yu N-N, Kärnefelt I, Thell A, Elix JA, Kim J, Kondratiuk AS and Hur J-S. 2014. A revised taxonomy for the subfamily Caloplacoideae (
Teloschistaceae, Ascomycota), based on molecular phylogeny. – Acta Botanica Hungarica 56: 93–123.
https://doi.org/10.1556/ABot.56.2014.1-2.10
Kondratyuk SY, Kärnefelt I, Thell A, Elix JA, Kim J, Kondratiuk AS and Hur J-S. 2015.
Brownlielloideae, a new subfamily in the
Teloschistaceae (Lecanoromycetes, Ascomycota). – Acta Botanica Hungarica 57: 321–341.
https://doi.org/10.1556/034.57.2015.3-4.6
Kondratyuk SY, Lőkös L, Halda JP, Roux C, Upreti DK, Schumm F, Mishra GK, Nayaka S, Farkas E, Park JS, Lee BG, Liu D, Woo J-J and Hur J-S. 2017. New and noteworthy lichen-forming and lichenicolous fungi 6. – Acta Botanica Hungarica 59: 137–260. https://doi.org/10.1556/034.59.2017.1-2.7
Kondratyuk SY, Lőkös L, Halda JP, Farkas E, Upreti DK, Thell A, Woo JJ, Oh SO and Hur JS. 2018. New and noteworthy lichen-forming and lichenicolous fungi 7. – Acta Botanica Hungarica 60: 115–184.
https://doi.org/10.1556/034.60.2018.1-2.8
Malcolm WM andGalloway DJ. 1997. New Zealand Lichens. Checklist, key, and glossary. – Museum of New Zealand, Te Papa Tongarewa, Wellington.
Mayrhofer H, Plattner K, Breuss O, Knudsen K, Sohrabi M, Rafiqpoor M D and Breckle S-W. 2023. The lichenized and lichenicolous fungi of Afghanistan. Plant and Fungal Systematics, 68(2), 440–461.
https://doi.org/10.35535/pfsyst-2023-0026
Memariani F. 2020. The Khorassan-Kopet Dagh Mountains. – In: Noroozi, J. (ed.), Plant Biogeography and Vegetation of High Mountains of Central and South West Asia. Springer Nature Switzerland AG, Cham, pp. 93–116. https://doi.org/10.1007/978-3-030-45212-4_3
Memariani, F, Zarrinpour V and Akhani H. 2016a. A review of plant diversity, vegetation, and phytogeography of the Khorassan-Kopet Dagh floristic province in the Irano-Turanian region (northeastern Iran–southern Turkmenistan).
Phytotaxa 249: 8–30.
https://doi.org/10.11646/phytotaxa.249.1.4
Memariani F, Akhani H and Joharchi MR. 2016b. Endemic plants of the Khorassan-Kopet Dagh floristic province in the Irano-Turanian region: diversity, distribution patterns and conservation status. Phytotaxa 249: 31–117. https://doi.org/10.11646/phytotaxa.249.1.5
Nakhaei M and Zokaei M. 2009. Identification and investigation of antibacterial activity of five indigenous lichens collected from the suburbs of Mashhad under in vitro conditions. – Azad University Biological Sciences Quarterly 7: 27–33.
Nash III TH, Ryan BD, Gries Cand Bungartz F. 2002. Lichen Flora of the Greater Sonoran Desert Region. Lichens Unlimited, Arizona State University, Tempe, Arizona, USA.
Nash III TH, Gries C and Bungartz F. (eds). 2007. Lichen Flora of the Greater Sonoran Desert Region. Vol. III. Lichens Unlimited, Arizona State University, Tempe.
Noroozi J, Talebi A, Doost Mohammadi M, Manaf Zadeh S, Asgarpour Z and Schneeweiss GM. 2019. Endemic diversity and distribution of the Iranian vascular flora across phytogeographical regions, biodiversity hotspots and areas of endemism. – Scientific Reports 9: 12991. https://doi.org/10.1038/s41598-019-49417-1
Obermayer W. 2002. Management of a lichen herbarium. – In: Kranner, I., Beckett, R. and Varma, A. (eds.),
Protocols in Lichenology. Springer Berlin Heidelberg, Berlin, Heidelberg, pp. 507–523.
https://doi.org/10.1007/978-3-642-56359-1_29
Rosentreter R, and McCune B. 1996. Distribution and ecology of Teloschistes contortuplicatus in North America. Evansia, 13, 10–13. https://doi.org/10.5962/p.346068
Ryan BD, Bungartz F and Nash TH. 2002. Morphology and anatomy of the lichen thallus. – In: Nash, T.H., Ryan, B.D., Gries, C. & Bungartz, F. (eds), Lichen Flora of the Greater Sonoran Desert Region. Lichens Unlimited, Arizona State University, Tempe, pp. 8–23.
Sayyad A, Ejtehadi H and Haji Moniri M. 2016a. The lichen flora of Hezar Masjed area. – In: Abstract Book of the Second Scientific Research Conference on Biology and Horticulture in Iran, 22 February 2016, Tehran, Iran(Online version).
Sayyad A, Ejtehadi H and Haji Moniri M. 2016b. The list of some lichens identified in Hezar Masjed area, Razavi Khorasan. – In: Abstract Book of the Second Scientific Research Conference on Biology and Horticulture in Iran, 22 February 2016, Tehran, Iran (Online version).
Seaward MRD, Sipman HJM, Schultz M, Maassoumi AA, Haji Moniri Anbaran M and Sohrabi M. 2004. A preliminary lichen checklist for Iran. – Willdenowia 34: 543–576.
https://doi.org/10.3372/wi.34.34218
Seaward MRD, Sipman HJM and Sohrabi M. 2008. A revised checklist of lichenized, lichenicolous and allied fungi for Iran. – Sauteria 15: 459–520.
Smith CW, Aptroot A, Coppins BJ, Fletcher A, Gilbert OL, James PW and Wolseley P. (eds) 2009. The Lichens of Great Britain and Ireland. British Lichen Society, London.
Sohrabi M and Sipman HJM. 2007. Lichenized fungi of Golestan National Park (NE Iran). – Mycologia Balcanica 4: 87–92. https://doi.org/10.5281/zenodo.2547960
Sohrabi M, Sipman H, Toghranegar Zand Nejad Sattari T. 2010. A contribution to the lichenized mycota of Zanjan province, Iran. – Iranian Journal of Botany 16: 126–129.
Sohrabi M, Ghiyasi A, Bordbar F, Safavi SR, Aliabadi F and Sipman H. 2019. A checklist of lichenized fungi of Kerman province, SE, Iran. – Mycologia Iranica 6: 21–32. https://doi.org/10.22043/mi.2020.121089
Szatala S. 1940. Lichenes. – In: Rechinger, K.H., Baumgartner, J., Petrak, F. and Szatala, S., Ergebnisse einer botanischen Reise nach dem Iran, 1937. Annalen des Naturhistorischen Museums in Wien 50: 521–533.
Szatala S. 1957. Prodromus einer Flechtenflora des Irans. – Annales Historico-Naturales Musei Nationalis Hungarici 8: 101–154.
Takhtajan AL. 1986. Floristic Region of the World. University of California Press, Berkeley.
https://doi.org/10.1525/9780520406568
Temina M, Kondratyuk S, Zelenko S, Wasser SP and Nevo E. 2005. Lichen-forming, lichenicolous and allied fungi of Israel. – In: Wasser, S.P. & Nevo, E. (eds), Biodiversity of Cyanoprokaryotes, Algae and Fungi of Israel. A.R.A. Ganter Verlag K.-G, Ruggell, Liechtenstein, pp. 1–384.
Valikhani R and Haji Moniri M. 2013. An addition to the knowledge of lichenicolous fungi of Iran. – Iranian Journal of Botany 19: 104–106.
Van Den Boom PPG and Haji Moniri M. 2018. Notes on the lichen genus Lecania (Ramalinaceae) in Iran, with the description of a new Arthonia species (Arthoniaceae). – Nova Hedwigia 107: 407–421.
https://doi.org/10.1127/nova_hedwigia/2018/0479
Vondrák J, Šoun J, Vondraková O, Fryday AM, Khodosovtsev A and Davydov E. 2012. Absence of anthraquinone pigments is paraphyletic and a phylogenetically unreliable character in the Teloschistaceae. – Lichenologist 44: 401–418. https://doi.org/10.1017/S0024282911000843
Westberg M and Sohrabi M. 2012. A conspectus of the lichen genus Candelariella (Candelariaceae, Ascomycota) in Southwest Asia with emphasis on Iran. – Nova Hedwigia 95: 531–546. https://doi.org/10.1127/0029-5035/2012/0060
Zakeri Z, Gasparyan A and Aptroot A. 2016. A new corticolous Megaspora (Megasporaceae) species from Armenia. – Willdenowia 46: 245–251. https://doi.org/10.3372/wi.46.46205
Zokaei M. 2002. Study on identification and distribution of lichens (order Lecanorales) in Mashhad and surroundings. – Rostaniha 3(1): 1–11. [Farsi].
Zokaei M. 2003. Study on identification and distribution of rocky lichens in surrounding mountains of Mashhad. – Journal of Scientific Research, University of Isfahan 1: 141–152. [Farsi].
Zokaei M and Abedi F. 2000. Study, recognition and introduction of six species of rocky lichens in north-eastern highlands of Mashhad (Kardeh area). – Agricultural Sciences and Technology 14: 2–14. [Farsi].
Zokaei M and Khazai M. 1999. Introducing several lichen species from Kalat Naderi. – In: Abstracts, 8th Iranian Biological Congress, University of Razi, 2–5 September 1999, Kermanshah, Iran.